• Blasdell, K.R., Duong, V., Eloit, M., Chretien, F., Ly, S., Hul, V., Deubel, V., Morand, S.,Buchy, P., 2016. Evidence of human infection by a new mammarenavirus endemic to Southeastern Asia. Elife, 5, e13135.

  • Campbell, C.B., 1966. Taxonomic status of tree shrews. Science, 153, 436.

  • Chen, L., Gu, W., Liu, C., Wang, W., Li, N., Chen, Y., Lu, C., Sun, X., Han, Y., Kuang, D., Tong, P.,Dai, J., 2019a. Characteristics of the tree shrew gut virome. PLoS One, 14, e0212774.

  • Chen, L., Yin, L., Zhou, Q., Peng, P., Du, Y., Liu, L., Zhang, Y., Xue, C.,Cao, Y., 2019b. Epidemiological investigation of fowl adenovirus infections in poultry in China during 2015-2018. BMC Vet Res, 15, 271.

  • Chen, Y.M., Hu, S.J., Lin, X.D., Tian, J.H., Lv, J.X., Wang, M.R., Luo, X.Q., Pei, Y.Y., Hu, R.X., Song, Z.G., Holmes, E.C.,Zhang, Y.Z., 2023. Host traits shape virome composition and virus transmission in wild small mammals. Cell, 186, 4662-4675.e4612.

  • Chen, Z., Chen, A.Y., Cheng, F.,Qiu, J., 2010. Chipmunk parvovirus is distinct from members in the genus Erythrovirus of the family Parvoviridae. PLoS One, 5, e15113.

  • Coutinho, F.H., Silveira, C.B., Gregoracci, G.B., Thompson, C.C., Edwards, R.A., Brussaard, C.P.D., Dutilh, B.E.,Thompson, F.L., 2017. Marine viruses discovered via metagenomics shed light on viral strategies throughout the oceans. Nat Commun, 8, 15955.

  • Darai, G., Rosen, A., Scholz, J.,Gelderblom, H., 1983. Induction of generalized and lethal herpesvirus infection in the tree shrew by intrahepatic transfection of herpes simplex virus DNA. J Virol Methods, 7, 305-314.

  • Decaro, N., Mari, V., Larocca, V., Losurdo, M., Lanave, G., Lucente, M.S., Corrente, M., Catella, C., Bo, S., Elia, G., Torre, G., Grandolfo, E., Martella, V.,Buonavoglia, C., 2016. Molecular surveillance of traditional and emerging pathogens associated with canine infectious respiratory disease. Vet Microbiol, 192, 21-25.

  • Drummond, A.J.,Rambaut, A., 2007. BEAST: Bayesian evolutionary analysis by sampling trees. BMC Evol Biol, 7, 214.

  • Easton, A.J., Domachowske, J.B.,Rosenberg, H.F., 2004. Animal pneumoviruses: molecular genetics and pathogenesis. Clin Microbiol Rev, 17, 390-412.

  • Edwards, R.A.,Rohwer, F., 2005. Viral metagenomics. Nat Rev Microbiol, 3, 504-510.

  • Fan, Y., Huang, Z.Y., Cao, C.C., Chen, C.S., Chen, Y.X., Fan, D.D., He, J., Hou, H.L., Hu, L., Hu, X.T., et al., 2013. Genome of the Chinese tree shrew. Nat Commun, 4, 1426.

  • Feng, Y., Feng, Y.M., Lu, C., Han, Y., Liu, L., Sun, X., Dai, J.,Xia, X., 2017. Tree shrew, a potential animal model for hepatitis C, supports the infection and replication of HCV in vitro and in vivo. J Gen Virol, 98, 2069-2078.

  • Feng, Y., Kuang, G., Pan, Y., Wang, J., Yang, W., Wu, W.-C., Pan, H., Wang, J., Han, X., Yang, L., Xin, G.-Y., Shan, Y.-T., Yu Gou, Q.-., Liu, X., Guo, D., Liang, G., Holmes, E.C., Gao, Z.,Shi, M., 2024. Small mammals in a biodiversity hotspot harbor viruses of emergence risk. National Science Review, 10.1093/nsr/nwae463.

  • Fuchs, E., 2024. Tree shrews. The UFAW handbook on the care and management of laboratory and other research animals, 324-339.

  • Geoghegan, J.L.,Holmes, E.C., 2017. Predicting virus emergence amid evolutionary noise. Open Biol, 7, 170189.

  • Harrach, B., Tarjan, Z.L.,Benkő, M., 2019. Adenoviruses across the animal kingdom: a walk in the zoo. FEBS Lett, 593, 3660-3673.

  • He, W.T., Hou, X., Zhao, J., Sun, J., He, H., Si, W., Wang, J., Jiang, Z., Yan, Z., Xing, G., et al., 2022. Virome characterization of game animals in China reveals a spectrum of emerging pathogens. Cell, 185, 1117-1129.e1118.

  • Hoan, N.X., Huy, P.X., Sy, B.T., Meyer, C.G., Son, T.V., Binh, M.T., Giang, D.P., Tu Anh, D., Bock, C.T., Wang, B., Tong, H.V., Kremsner, P.G., Song, L.H., Toan, N.L.,Velavan, T.P., 2019. High Hepatitis E virus (HEV) Positivity Among Domestic Pigs and Risk of HEV Infection of Individuals Occupationally Exposed to Pigs and Pork Meat in Hanoi, Vietnam. Open Forum Infect Dis, 6, ofz306.

  • Jiang, L., Lu, C.,Sun, Q., 2021. Tree Shrew as a New Animal Model for the Study of Dengue Virus. Front Immunol, 12, 621164.

  • Johnson, M., Zaretskaya, I., Raytselis, Y., Merezhuk, Y., Mcginnis, S.,Madden, T.L., 2008. NCBI BLAST: a better web interface. Nucleic Acids Res, 36, W5-9.

  • Jones, K.E., Patel, N.G., Levy, M.A., Storeygard, A., Balk, D., Gittleman, J.L.,Daszak, P., 2008. Global trends in emerging infectious diseases. Nature, 451, 990-993.

  • Kane, Y., Tendu, A., Li, R., Chen, Y., Mastriani, E., Lan, J., Catherine Hughes, A., Berthet, N.,Wong, G., 2024. Viral diversity in wild and urban rodents of Yunnan Province, China. Emerg Microbes Infect, 13, 2290842.

  • Kearse, M., Moir, R., Wilson, A., Stones-Havas, S., Cheung, M., Sturrock, S., Buxton, S., Cooper, A., Markowitz, S., Duran, C., Thierer, T., Ashton, B., Meintjes, P.,Drummond, A., 2012. Geneious Basic: an integrated and extendable desktop software platform for the organization and analysis of sequence data. Bioinformatics, 28, 1647-1649.

  • Krishnamurthy, S.R.,Wang, D., 2017. Origins and challenges of viral dark matter. Virus Res, 239, 136-142.

  • Kumar, S., Stecher, G.,Tamura, K., 2016. MEGA7: Molecular Evolutionary Genetics Analysis Version 7.0 for Bigger Datasets. Mol Biol Evol, 33, 1870-1874.

  • Letko, M., Seifert, S.N., Olival, K.J., Plowright, R.K.,Munster, V.J., 2020. Bat-borne virus diversity, spillover and emergence. Nat Rev Microbiol, 18, 461-471.

  • Liu, P., Qiu, Y., Xing, C., Zhou, J.H., Yang, W.H., Wang, Q., Li, J.Y., Han, X., Zhang, Y.Z.,Ge, X.Y., 2019. Detection and genome characterization of two novel papillomaviruses and a novel polyomavirus in tree shrew (Tupaia belangeri chinensis) in China. Virol J, 16, 35.

  • Lole, K.S., Bollinger, R.C., Paranjape, R.S., Gadkari, D., Kulkarni, S.S., Novak, N.G., Ingersoll, R., Sheppard, H.W.,Ray, S.C., 1999. Full-length human immunodeficiency virus type 1 genomes from subtype C-infected seroconverters in India, with evidence of intersubtype recombination. J Virol, 73, 152-160.

  • Luis, A.D., Hayman, D.T., O'shea, T.J., Cryan, P.M., Gilbert, A.T., Pulliam, J.R., Mills, J.N., Timonin, M.E., Willis, C.K., Cunningham, A.A., Fooks, A.R., Rupprecht, C.E., Wood, J.L.,Webb, C.T., 2013. A comparison of bats and rodents as reservoirs of zoonotic viruses: are bats special? Proc Biol Sci, 280, 20122753.

  • Meerburg, B.G., Singleton, G.R.,Kijlstra, A., 2009. Rodent-borne diseases and their risks for public health. Crit Rev Microbiol, 35, 221-270.

  • Mokili, J.L., Rohwer, F.,Dutilh, B.E., 2012. Metagenomics and future perspectives in virus discovery. Curr Opin Virol, 2, 63-77.

  • Nie, F.Y., Tian, J.H., Lin, X.D., Yu, B., Xing, J.G., Cao, J.H., Holmes, E.C., Ma, R.Z.,Zhang, Y.Z., 2019. Discovery of a highly divergent hepadnavirus in shrews from China. Virology, 531, 162-170.

  • Ren, S., Wang, C., Zhang, X., Zhao, L., Wang, X., Yao, W., Han, Q., Wang, Y., Fan, M., Gao, X., Xiao, S., Wang, X.,Yang, Z., 2017. Phylogenetic and pathogenic characterization of a pigeon paramyxovirus type 1 isolate reveals cross-species transmission and potential outbreak risks in the northwest region of China. Arch Virol, 162, 2755-2767.

  • Sayers, E.W., Bolton, E.E., Brister, J.R., Canese, K., Chan, J., Comeau, D.C., Connor, R., Funk, K., Kelly, C., Kim, S., et al., 2022. Database resources of the national center for biotechnology information. Nucleic Acids Res, 50, D20-d26.

  • Schondorf, E., Bahr, U., Handermann, M.,Darai, G., 2003. Characterization of the complete genome of the Tupaia (tree shrew) adenovirus. J Virol, 77, 4345-4356.

  • Song, X., Li, Y., Huang, J., Cao, H., Zhou, Q., Sha, X.,Zhang, B., 2021. An emerging orthopneumovirus detected from dogs with canine infectious respiratory disease in China. Transbound Emerg Dis, 68, 3217-3221.

  • Sullivan, M.B., 2015. Viromes, not gene markers, for studying double-stranded DNA virus communities. J Virol, 89, 2459-2461.

  • Temmam, S., Davoust, B., Berenger, J.M., Raoult, D.,Desnues, C., 2014. Viral metagenomics on animals as a tool for the detection of zoonoses prior to human infection? Int J Mol Sci, 15, 10377-10397.

  • Tidona, C.A., Kurz, H.W., Gelderblom, H.R.,Darai, G., 1999. Isolation and molecular characterization of a novel cytopathogenic paramyxovirus from tree shrews. Virology, 258, 425-434.

  • Watsa, M., 2020. Rigorous wildlife disease surveillance. Science, 369, 145-147.

  • Xiong, Y.Q., Mo, Y., Chen, M.J., Cai, W., He, W.Q.,Chen, Q., 2018. Detection and phylogenetic analysis of torque teno virus (TTV) carried by murine rodents and house shrews in China. Virology, 516, 189-195.

  • Xu, L., Yu, D.D., Ma, Y.H., Yao, Y.L., Luo, R.H., Feng, X.L., Cai, H.R., Han, J.B., Wang, X.H., Li, M.H., Ke, C.W., Zheng, Y.T.,Yao, Y.G., 2020. COVID-19-like symptoms observed in Chinese tree shrews infected with SARS-CoV-2. Zool Res, 41, 517-526.

  • Yang, Z.F., Zhao, J., Zhu, Y.T., Wang, Y.T., Liu, R., Zhao, S.S., Li, R.F., Yang, C.G., Li, J.Q.,Zhong, N.S., 2013. The tree shrew provides a useful alternative model for the study of influenza H1N1 virus. Virol J, 10, 111.

  • Zhao, F., Guo, X., Wang, Y., Liu, J., Lee, W.H.,Zhang, Y., 2014. Drug target mining and analysis of the Chinese tree shrew for pharmacological testing. PLoS One, 9, e104191.

  • Zhou, H., Tian, R.R., Wang, X.R., Yang, J.X., Wang, Y.X., Zhao, M.L., Zhang, X.D., Ma, Y.H., Lv, L.B., Holmes, E.C., Zheng, Y.T.,Shi, W.F., 2024. Identification of novel mammalian viruses in tree shrews (Tupaia belangeri chinensis). Zool Res, 45, 429-438.